Rome, 1943: University student Marina Tozzi is on her way home when she finds out that her father has been killed for harboring a Jewish artist in their home. Fearful of the consequences, Marina flees to Villa I Tatti, the Florence villa of her father's American friend Bernard Berenson and his partner Belle da Costa Greene, the famed librarian who once curated J.P. Morgan's library. Florence is a hotbed of activity as partisans and Germans fight for control of the city. Marina, an art expert, begins helping Bernard catalog his library as he makes the difficult trek to neutral Switzerland, helping to hide precious cultural artifacts from the Germans. Adding to the tension, their young neighbor Carlos, a partisan, seeks out Marina for both her art expertise and her charm. Marina, swept up in the romance, dreams of a life together after the war. But when Carlos disappears, all of Marina's assumptions about her life in Florence are thrown into doubt, and she'll have to travel halfway around the world to unravel what really happened during the war.